E. J.  Tizado, Núñez-Pérez, E., and Salgado, J. M., ?Morphometric relationships among populations of the cave-dwelling beetle Quaestus nietoi (Salgado, 1988) (Coleoptera: Leiodidae)?, ASef pttts (n. s.), vol. 33, pp. 215-231., 1997.