in: T. Nakane, K. Ohbayashi, S. Nomura, Y. Kurosawa, Iconographia insectorum japonicum Colore naturali edita, volume 2, Coleoptera

Scratchpads developed and conceived by (alphabetical): Ed Baker, Katherine Bouton Alice Heaton Dimitris Koureas, Laurence Livermore, Dave Roberts, Simon Rycroft, Ben Scott, Vince Smith